Amphisphaeriales Hawksw. & O.E. Erikss., Syst. Ascom. 5(1): 177 (1986)

Amphisphaeriales has been accepted in consecutive studies using multigene phylogenies in the subclass Xylariomycetidae (Senanayake et al. 2015, Samarakoon et al. 2016b, Hongsanan et al. 2017, Crous et al. 2018b). However, based on previous studies, Wijayawardene et al. (2018a) accepted 11 families in Amphisphaeriales, i.e. Amphisphaeriaceae, Apiosporaceae, Beltraniaceae, Clypeophysalosporaceae, Hyponectriaceae, Melogrammataceae, Oxydothidaceae, Phlogicylindriaceae, Pseudomassariaceae, Sporocadaceae and Vialaeaceae, while Cainiaceae, Coniocessiaceae, and Iodosphaeriaceae were accepted in Xylariomycetidae families incertae sedis. In this study, Amphisphaeriales is a moderately supported clade sister to Xylariales and comprises 17 families viz. Amphisphaeriaceae, Apiosporaceae, Beltraniaceae, Castanediellaceae, Clypeophysalosporaceae, Cylindriaceae, Hyponectriaceae, Iodosphaeriaceae, Melogrammataceae, Oxydothidaceae, Phlogicylindriaceae, Pseudomassariaceae, Pseudosporidesmiaceae, Pseudotruncatellaceae, Sporocadaceae, Vialaeaceae, and Xyladictyochaetaceae. The divergence time for Amphisphaeriales has been estimated as 133 MYA. This number did not include Vialaeaceae because this family goes to Xylariales in MCC tree (Fig. 2). Currently there are 17 families and 88 genera in this order (this paper). 


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The webpage Sordariomycetes provides an up-to-date classification and account of all genera of the class Sordariomycetes.


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